Origin of anatomically modern humans

How evolution proceeds and species arise are affected by many different processes. But, if members of every generation of the clade that led from the probable common ancestor of ourselves, Neanderthals, Denisovans and other hominins of the last 700 ka or so – widely thought to have been Homo heidelbergensis­ – were found as perfectly preserved fossils they would show gradually shifting anatomical features that from time to time and place to place would diverge to lead to different species. If, also, every specimen was accurately dated then there would be the last part of the human evolutionary bush laid out in a 3-D graphic. That is never going to be possible, of course. Human fossils are rare and there are few of them that are well-preserved. So the field of human origins throws up surprises on a regular basis, and if palaeoanthropologists were more dogmatic than most of them actually are there would be equally regular, public displays of the eating of hats.

As regards early modern H. sapiens, fossils from a couple of sites in Ethiopia have been the oldest known, at between 160 to 195 ka, for the last 15 years. However, in the 1960s quarry workers at Jebel Irhoud in SW Morocco exposed the infill of a cave network in which were found numerous items of the Levallois stone-tool technology, some human bone fragments that included a brain case and many dismembered and cut bones of prey animals. Initially they were thought to date from about 40 ka and to represent an African form of Neanderthals. Subsequently, re-evaluation of the remains revealed a greater likelihood that they were from modern humans, but too young to be of great interest. An upgraded date of ~160 ka caused them to be considered  as peripheral to the core group of Ethiopian early modern humans. DNA analyses then suggested modern humans to have split from Neanderthals about 500 ka ago. Members of the French-Moroccan team that did the original work, accompanied by other scientists, recently re-excavated the site and exhumed a much richer fossil haul that pin-pointed an anatomically modern human (AMH) provenance, albeit with some archaic characteristics (Hublin, J.-J. and 10 others, 2017. New fossils from Jebel Irhoud, Morocco and the pan-African origin of Homo sapiens. Nature, v. 546, p. 289-294; doi:10.1038/nature22336), which can be referred to as ‘pre-modern’ H. sapiens. The bombshell stemming from their work was the precise dating of the fossils and their stratigraphic context by other members of the team (Richter, D. and 11 others. The age of the hominin fossils from Jebel Irhoud, Morocco, and the origins of the Middle Stone Age. Nature, v. 546, p. 293-296; doi:10.1038/nature22335), which yielded 315±34 ka from fire-heated flint fragments and 286±32 ka from a human tooth. Both dates are far older than the previously accepted maximum of 200 ka for AMH.

The early evolution of fully modern humans seems to have spanned the whole of Africa, rather than being set in an Ethiopian heartland, a view partly supported by a fragmentary 260 ka fossil from South Africa bearing close resemblance to the Moroccan individuals. Interestingly, Levallois stone tools, as their name suggests, are widespread in both Africa and Europe at around 300 ka, although that is not proof that AMH migrated out of Africa around 300 ka, for Neanderthals may also have been using a similar flint flaking method (another space to be watched).

See also:  Stringer C. & Galway-Witham, J., 2017. On the origin of our species.  Nature, v. 546, p. 212-215; doi:10.1038/nature 546212a.

You can find more information on migration of modern humans here.

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